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Travelers using the Elizabeth line could save money on their journey in and out of certain stations.
Anyone traveling to the city center from outside the capital, such as Reading or Shenfield, could save money due to a ticket sale gap.
A rush hour journey from Reading to Tottenham Court Road costs £ 28.50 for an adult entering and leaving with a contactless card, according to Transport for London fare search engine.
However, a trip from Reading to Slough costs just £ 5.90 while Slough on Tottenham Court Road costs £ 11.80.
The rift exists as prices shift between Transport for London (TfL) fares and National Rail fares to the outermost stations in the capital, according to the Telegraph.
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This allows travelers to save £ 10.80 if they choose to enter and exit Slough instead of traveling directly from Reading.
Across the line, a peak hourly fare from Shenfield to Paddington costs £ 13, but travelers could save money by splitting the trip.
Traveling from Shenfield to Harold Wood costs £ 3.60, while Harold Wood to Paddington costs £ 5.50, saving £ 3.90.
Norman Baker of Campaign for Better Transport, a campaign group, told the Telegraph that the system was “complicated” and “confusing” and said a “root and branch reform” of the entire rail system was needed.
Meanwhile, the first trains on the Elizabeth line ran on Tuesday morning as Mayor Sadiq Khan praised a “historic day” for the capital.
The first train left Abbey Wood Station at 6.30am, followed three minutes later by a similar eastbound service from Paddington.
About 130,000 people traveled on the line in the early hours of Tuesday morning, according to TfL figures. The line runs from Reading in Berkshire and Heathrow Airport in west London to Abbey Wood in south east London and Shenfield in Essex.
The £ 20bn project is the largest upgrade to the capital’s transport infrastructure in more than 50 years.
A TfL spokesman said: “The fares between Reading and Slough are set by GWR and remain the same as those operating under TfL Rail.
“As with all rail travel across the UK, there will be times when customers will be able to benefit from ticket distribution or pay fares as they go through the departure and return stations with the your contactless card.
“Our usage-based payment system calculates the most appropriate fare for your trip, including whether a daily or weekly limit would apply. Contactless payment can be used across the entire Elizabeth line.”